Application Information
Applications for the 2026–2027 academic year to BBS are now closed. Applications for the 2027–2028 academic year will open in early fall 2026, with an anticipated deadline of December 1, 2026.
Welcome!
Biological and Biomedical Sciences (BBS) at Harvard University is a full-time Ph.D. training program spanning a very broad range of topics in modern life science research. Our research areas extend from angstrom-level atomic resolution across the spectrum to whole organismal animal physiology and behavior. Our scientists use a huge array of systems from prokaryotic and viral organisms, to invertebrate models, to many different vertebrate species. While we are physically based at Harvard Medical School (HMS), we are also part of the Harvard Integrated Life Sciences (HILS) federation, and thus our students have access to hundreds of thesis research advisors across the Harvard landscape. As the largest of Harvard’s PhD programs, BBS students currently work in a total of 172 laboratories within our network of Harvard schools and affiliated hospitals. BBS is guided by leadership from a consortium of basic science departments at HMS, including Biological Chemistry and Molecular Pharmacology, Cell Biology, Genetics, Microbiology, and Stem Cell and Regenerative Biology.
BBS Community
Because BBS students pursue thesis research in a diverse array of topics, model systems and laboratories, the potential for interdisciplinary collaboration, co-mentorship and collegial cross-pollination is one of the features that draws many students to our program. Our BBS community brings scholars from all over the world (currently 32 different countries), and a myriad of different backgrounds and perspectives. The scale and scope of BBS is also a key reason that we treasure a sense of intellectual and social community. For this reason, we encourage and support student-led initiatives, as well as multiple forms of service and leadership. In the admissions process, we look for candidates who have demonstrated their passion for community and service in the recent past through a range of activities and contributions to their own communities; the part of the application where we ask students to describe these accomplishments is the Personal Statement.
Preparation and Prerequisites
Although BBS does not have strict curricular prerequisites, as a program founded on understanding fundamental biological and molecular mechanisms, we believe that students who have course backgrounds that include a combination of biochemistry, cell biology, genetics and molecular biology, will find our graduate level curriculum most appropriate in level and scope. We prefer students to have prior course experience with statistical methods, and often find that some familiarity with computational programming is extremely helpful. Equally important as curricular background, we feel that prior research experiences are vital to the preparation of candidates to enter our program. For this reason, it is important that applicants articulate their research experiences clearly and succinctly in their Statement of Purpose.
What we are looking for in candidates
There are many ingredients that contribute to a scientist’s success in leading the transformative breakthroughs that bring deeper understanding of biological systems and phenomena. For this reason, our review of each application is holistic – weighing every element of the candidate’s portfolio and actual experiences as described and documented in several areas of the application. We actively search for candidates that have shown a true love for the process of discovery and problem-solving. We value creativity, resourcefulness and intellectual independence. We recognize that commitment, dedication and hard work are essential to successful scientific investigation. However, we also appreciate that research almost always presents many obstacles. For this reason, we value persistence and resilience in the face of adversity. The pursuit of doctoral training is a path to professional development. It requires competence in a variety of areas, including scholarly mastery of many topics and technologies, diligent application of these skills in the laboratory, and the ability to communicate effectively with others inside and outside of the field.
Review Process
Due to our large size and intellectual breadth, BBS organizes its admissions process in a way analogous to NIH scientific review. BBS faculty reviewers are assigned to one of four subcommittees: Bacteriology, bacterial pathogenesis, and the microbiome; Cancer Biology and Therapeutics; Genetics and Genomics; and Molecular, Cellular and Developmental Mechanisms. Applicants are asked to select one of our subcommittees that best aligns with their own planned area of future research training and prior background. All subcommittees use the same overall process and criteria, however, readers in each committee bring domain-relevant expertise to the review process. Importantly, by grouping faculty in smaller review cohorts, discussions about each candidate portfolio can be more detailed and engaged, with greater focus on each applicant’s potential for a creative and productive research career.
Although our admissions process has four streams of review, this in no way restricts or commits our applicants to a specific area of thesis research once they begin graduate study. BBS remains a completely open umbrella program that allows students the freedom to grow and change as they navigate our first-year curriculum and laboratory rotations. While BBS and its supporting departments and institutions provide different affinity groups and activities to foster intellectual community for different areas of science, we are highly committed to innovative and interdisciplinary research that connects various fields of study within our extensive community of scholars.
BBS Sub-Committees
Molecular, Cellular and Developmental Mechanisms
The Molecular, Cellular and Developmental Mechanisms subcommittee brings together researchers seeking to understand form and function in biology at many scales. Faculty investigate molecular machines that control cell signaling, gene expression, cell mechanics, quality control and metabolism, as well as the processes by which the body’s organelles, cells and tissues are formed, maintained and repaired. Faculty also use these fundamental insights to inform how genetic alterations drive disease and to transform that understanding into effective therapies.
Genetics and Genomics
The Genetics and Genomics committee is composed of faculty who use genetic and genomic approaches in all the major model systems, including humans, to explore basic mechanisms of biology and investigate how these mechanisms relate to disease in people.
Bacteriology, bacterial pathogenesis, and the microbiome
Faculty members serving on the Bacteriology, bacterial pathogenesis, and the microbiome subcommittee have broad expertise in fundamental mechanisms of bacterial cell and molecular biology, bacterial pathogenesis, and the microbiome.
Cancer Biology and Therapeutics
Faculty members serving on the Cancer Biology and Therapeutics have broad expertise in deconstructing basic biological and cellular mechanisms of cancer development, cancer genetics and epigenetics, cancer immunology, and all facets of therapeutic development.
Interviews
After thoroughly reading and appreciating the many features of each application, our admissions team will select a subset of candidates for further interview. Although we learn much from each application, we find that active conversations provide us with a very helpful way to confirm and build our emerging picture of each candidate’s unique qualities and the depth of their experiences. We also find that interviews help our candidates understand much more about our program and our faculty.
Requirements
Minimal requirements include a bachelor’s degree and undergraduate preparation in the sciences. Given our holistic admissions process, it is not possible to provide feedback on any application for admission prior to review by the admissions committees.
- Online application form
- Unofficial transcripts uploaded to the application
- Three Letters of recommendation
- Statement of Purpose (SoP)
- Personal Statement
- Demonstration of English proficiency (TOEFL or IELTS), where required
- Curriculum Vitae (CV) or Resume
- Application fee payment ($105)*
*Application Fee Waiver: we are committed to ensuring that the application fee does not create a financial obstacle. Applicants can determine eligibility for a fee waiver by completing a series of questions in the application. Once these questions have been answered, the application system will provide an immediate response regarding fee waiver eligibility. Please note - the program is not able to directly authorize fee waivers.
Admissions Timeline
Application Portal Opens | Early September |
Application Deadline | December 1, 5:00 PM EST |
Interview Invitations Sent | Mid-December |
Interview Weekends | January 21 - 25, 2027 & February 4 - 7, 2027 |
Admit & Waitlist Offers Sent | Mid-February |
Decision Deadline | April 15th |
How to Apply
The application will open in early September. All materials must be submitted through the Harvard Griffin GSAS Applicant Portal by 5:00 PM (EST) on December 1.
Selecting the Degree Program: On the GSAS application, select "Biological and Biomedical Sciences" as your Degree Program.
Selecting BBS Subcommittee: Please review the descriptions of our four subcommittees and select the one that best corresponds to your intended area of future research as well as your previous academic and professional experience. Choosing the subcommittee most closely aligned with your interests and background will ensure that your application is evaluated by experts who are best suited to assess your potential and fit for the program. If you are unsure which subcommittee to choose, we recommend considering both your proposed research direction and your foundational training to make the most informed decision. Please note - If you choose "Molecular, Cellular and Developmental Mechanisms" as your preferred area of interest, please further specify your focus by selecting one of the following options: a) Molecular Mechanisms, b) Structural Biology, c) Cellular Biology, or d) Stem Cell Biology and Development. This selection will help us direct your application to the most relevant reviewers, but it does not commit you to a specific area.
Faculty of interest: Since we ask students to indicate one of the general areas represented in our admissions committee, we don’t ask students to list specific faculty of interest in the application. Once candidates are invited for interviews, however, we will inquire if there are particular faculty that each applicant may wish to meet, if available. Students are not required to contact faculty or secure a thesis laboratory prior to submitting their applications to BBS. Once matriculated, students will have the opportunity to rotate in labs prior to declaring a thesis mentor at the end of their G1 year (July 1st).
Statement of Purpose (SoP): Through the SoP, we’re looking to get to know you as a scientist. How did your background help shape your interests and experiences in science? What are your goals for graduate school? How do your research experiences fit within this context? You are welcome to briefly mention multiple research experiences as they relate to your trajectory, but your SoP should focus on your primary research experience, discussing the question/topic that drove the project, its importance, your hypotheses, how you tested your hypotheses, your findings or possible outcomes, and how you interpreted those outcomes. In this part of the application, we're eager to see how you demonstrate your ability to approach problems and ideas with a scientific mindset. (Not to exceed 1,000 words)
Ultimately, the statement should provide evidence of research readiness through descriptions of past projects with clarity including explaining your role, methods, and what was learned.
Personal Statement: scientific discovery and professional development benefit greatly from the exchange of ideas and perspectives within the community of scholars. This short essay is a dedicated space for applicants to describe the specific prior activities, leadership roles, service contributions and lived experiences that illustrate their unique perspectives, their values, and their commitment to intellectual community. It is important that applicants do not use this essay as space to merely expand on the research experiences discussed in the Statement of Purpose, but rather, use the Personal Statement to describe additional dimensions of their aspirations, experiences and accomplishments that will guide their future contributions to the research community around them. (Not to exceed 500 words)
Letters of Recommendation: applicants are required to submit three letters of recommendation. We rely heavily on sincere, candid and detailed accounts of each candidate’s accomplishments in their scholarly pursuits and research training from mentors and advisors who have taken the time to cultivate a deep familiarity and understanding of each candidate’s strengths and unique attributes.
Transcripts: applicants are required to upload unofficial transcripts from each institution attended. We do not accept transcripts received by mail or email during the application submission and review process. Official transcripts are only required from those who have been admitted and accepted the offer of admission. Transcripts from international institutions must be in English or be accompanied by a certified English translation. All translations must be literal and complete versions of the original records. For more information, please visit the GSAS Transcripts Page.
GRE: BBS no longer accepts GRE scores. If you submit scores, they will not be seen by our admissions committees.
TOEFL/IELTS: if English is not your native language and you completed your undergraduate degree in a language other than English, you are required to demonstrate English proficiency by submitting TOEFL (minimum of 100) or IELTS Academic (7.0) scores. In addition, candidates will need to demonstrate a minimum speaking subsection test score of 23 (old scale)/4.5 (new scale) for TOEFL or a minimum speaking subsection test score of 6.5 for IELTS. Applicants who took tests more than two years ago (i.e., before January 5, 2025, for Fall 2027 admission) must retake them. We are unable to accept any other proof of proficiency. A master’s degree or other graduate degree is not accepted as proof of English proficiency. This requirement applies to all applicants regardless of residency or citizenship status. If you received an undergraduate degree from an institution in the United States in which English was not the primary language of instruction but a significant portion of your program was conducted in English, you may petition for a waiver of the TOEFL/IELTS requirement. Please note this ONLY applies to US institutions. Contact the office of admissions for details.
HILS
Although students may apply to multiple programs, they are strongly encouraged to explore the options carefully and focus on those that best match their interests and goals. Below is an illustration of the Harvard HILS PhD landscape.
Financial Support
Tuition & Stipend
BBS students receive full tuition and stipend support, as outlined in their financial aid offer, while they are enrolled and making satisfactory progress toward the Ph.D. degree.
Health Insurance and Benefits
Students are eligible to enroll in the Harvard University Student Health Program (HUSHP) and are not responsible for paying the yearly health fee. Click the link for a summary of the benefits offered with HUSHP. Please refer to their website to learn more about these benefits.
Relocation Allowance & Other Benefits
Upon matriculation to BBS, all students are given a one-time $1,000 relocation allowance. For additional benefits, please visit the OGE Financial Support page.
BBS does not discriminate against applicants or students on the basis of race, color, national origin, ancestry, or any other protected classification.